Random thing I just noticed: the Serpentine Generals have glowing eyes where the regular Serpentine in their tribes do not.
Also, on another note, Pythor does as well:
That, plus the tail and his possession of the Anacondrai staff, leads me to believe that back during the production and airing of Season 1, he may have been meant to be the sole General of the Anacondrai before Season 4 made it canon that the Anacondrai had not one, but multiple generals.
I wish the Ninjago kept on animating actual facial expressions for Skales (instead of just a constant rbf) after season 2 because it gave him so much more personality. (It also gives funky screenshots like these)
Something I haven't seen brought up: Skales helped burn down the original monastery, but he also bothered to attend Nya's memorial in Seabound. Having a kid really seemed to have changed Skales for the better and I kinda love it.